sábado, 11 de agosto de 2007

Fisiología.

Fisiología celular, Meiosis y Mitosis
Fisiología celular
1) RELACION: Esta función permite la interacción con el medio ambiente, y se basa en movimientos internos (ciclosis) o externos (tropismos, taxismos).
Ciclosis:
Movimiento circulatorio que se produce en el citoplasma por cambios de estado y por acción del citoesqueleto ante estímulos externos.
Tropismos: Son movimientos de orientación en el crecimiento de las
células vegetales hacia o en contra de un estímulo externo (Ej: fototropismo positivo en hojas y negativo en raíces).
Taxismos: Son movimientos de traslación de
células animales producido por cilias, flagelos o ameboidales como respuesta a estímulos.
2) REPRODUCCIÓN: Es la
propiedad de engendrar organismos similares o iguales asegurando la supervivencia de la especie. Puede ser por mitosis (la célula madre origina 2 células con igual número de cromosomas) o por meiosis (la célula madre origina 4 células con la mitad del número cromosómico).
3)
NUTRICIÓN: Es un conjunto de funciones para obtener materia y energía por intercambio con el ambiente. En heterótrofos, las funciones son: ingestión, digestión, asimilación, excreción, respiración y circulación. En autótrofos, son: fotosíntesis, respiración y circulación.
Heterótrofos:
Ingestión:
La célula incorpora materia por endocitosis, y se forma una vacuola alimenticia.
Digestión: Un lisosoma primario se acerca a la vacuola alimenticia, se fusionan sus membranas, y se forma un lisosoma secundario. Allí las
enzimas digestivas desdoblan las moléculas complejas en simples.
Circulación: Por la digestión, las
proteínas se desdoblan en aminoácidos, los lípidos en ácidos grasos y los hidratos de carbono en monosacáridos. Las moléculas simples ya pueden ser asimiladas, y para ello deben circular por medio de la ciclosis.
Excreción: Las sustancias no asimilables se acumulan en vacuolas o se fusionan con la membrana plasmática, y por exocitosis expulsan su contenido.
Respiración: Se produce gracias a la
materia y energía obtenidas de los alimentos digeridos. Es el proceso por el cual la glucosa es oxidada CO2 y H2O en presencia de O2, con liberación de energía. Comprende 3 etapas:
Glucósis: Se realiza en el citoplasma donde hay
enzimas que degradan parcialmente la glucosa, liberando energía (ATP).
Ciclo de Krebs: Ocurre en la
matriz mitocondrial por una acción enzimática. Se produce liberación de CO2 y energía.
Cadena respiratoria: Se produce en las crestas mitocondriales donde hay
enzimas que forman la cadena respiratoria. Finalmente, la glucosa es degradada totalmente.
Autótrofos:
Fotosíntesis: Los vegetales elaboran glucosa a partir de
agua, sales CO2 y energía luminosa captada por la clorofila. Los cloroplastos están formados por tres membranas los tilacoides se apilan formando granas dentro de la matriz, y la clorofila está en la superficie interna de los tilacoides. La fotosíntesis se realiza en el parénquima clorofiliano de las plantas y consta de 2 etapas: lumínica (se realiza en los tilacoides en presencia de luz) y oscura (no necesita luz y ocurre en la matriz).
Fase lumínica: La energía lumínica es captada por la clorofila y transformada en energía
química. La energía química se almacena en compuestos como el ADP que al incorporar energía se transforma en ATP. La energía del ATP se utiliza para romper la molécula de agua y separarla en H2 y O2, proceso de hidrólisis. El O2 sale por los estomas y el H2 queda detenido en un compuesto que actúa como aceptor de H2.
Fase oscura: Se utiliza la energía acumulada en el ATP, el cual cede un ácido fosfórico y origina ADP, liberando energía. Los aceptores ceden el H2 que se combina con el CO2 usando energía del ATP. Esa combinación origina glucosa. Este
proceso se llama ciclo de Calvin. A partir de la glucosa se originan azúcares (almidón y sacarosa) o lípidos (que se acumulan en oleoplastos) o proteínas (en proteoplastos). El transporte de estas sustancias se realiza por el floema.
Circulación: Responde a la
teoría tenso-ccheso-transpiratoria. El agua entra en la raíz por ósmosis, atraviesa la epidermis (rizodermis), pasa al apénquima cortical, y luego entra en el xilema, que se encargará de distribuir el agua las sales a toda la planta. Para que el agua ascienda requiere de cohesión de sus moléculas que se unen formando columnas, las cuales permanecen unidas e todo su recorrido por los vasos del xilema. Cuando la planta transpira por los estomas, se genera un vacío temporario en los vasos xilemáticos que sufren una tensión que hacen ascender la columna de agua. El floema es otro tejido conductor compuesto por células vivas y paralelo al xilema, que transporta la glucosa desde la hoja hasta el resto del vegetal (camino adverso del xilema).

Mitosis
Es la división celular que consiste en que a partir de una
célula se obtienen 2 células hijas, genéticamente idénticas a la madre. Se produce en cualquier célula eucarionte, ya sea diploide o haploide y como mantiene invariable el número de cromosomas, las células hijas resultarán diploides, si la madre era diploide o alploide. La división del citoplasma se llama citocinesis, y la división del núcleo, cariocinesis. Algunas células no realizan mitosis y permanencen en un estado interfásico, pero otras la realizan frecuentemente (células embrionarias, células de zonas de crecimiento, células de tejidos sujetos a desgaste.).
Función: crecimiento y
desarrollo del organismo multicelular, y la regeneración de tejidos expuestos a destrucción de células. En unicelulares, cumple la función de reproducción asexual.
Cada
mitosis está precedida por una interfase, donde se produce la duplicación del material genético. Actúa como un mecanismo que asegura que cada célula hija reciba la misma información genética.
Etapas: Profase, Prometafase, Metafase, Anafase y Telofase.
PROFASE: La cromatina se condensa para formar los
cromosomas y los 2 centríolos migran a polos opuestos organizando un sistema de microtóbulos (aparato mitótico) para permitir la migración de los cromosomas. El aparato mitótico está constituído por:
Centríolos: Están rodeadas por el centrosoma. A medida que cada centríolo migra, tiene un hijo y cuando llega al polo se ven 2.
Ásteres: Conjunto de microtóbulos cortos que se extienden desde cada centríolo.
Huso acromático: Tiene forma de ovoide y formado por muchos microtóbulos sin ramificaciones.
Cada cromosoma está constituido por 2 cromátidas unidas por el centrómero. La envoltura nuclear se desorganiza y sus fragmentos no se distinguen del retículo endoplasmático. Desaparece el nucleolo.
PROMETAFASE: Los
cromosomas condensados migran hacia la placa ecuatorial del huso acromático.
METAFASE: Los cromosomas se alínean en el plano ecuatorial, y cada uno están unido por su centrómero a una fibra del huso acromático.
ANAFASE: Las 2 cromátides de cada cromosoma se separan por fisión del centrómero y se dirigen hacia polos opuestos. El
movimiento de los cromosomas hijos hacia los polos se debe a un acortamiento de las fibras cromosómicas y se alargan las fibras interzonales.
TELOFASE: El huso mitótico y los ásteres se desorganizan. Alrededor de cada
grupo cromosómico se organiza una envoltura nuclear a partir del re´ticulo endoplasmático y de la envoltura original. Los cromosomas se dispersan y retoman el aspecto de cromatina que tenían antes de iniciarse la división. Los nucleolos reaparecen a partir de sus organizadores.
Citocinesis:
La división del citoplasma se produce junto con la telofase. Se produce un surco en la membrana plasmática, producidom por un anillo de mocrofilamentos unidos a ella. Las 2 células hijas se serparan, distribuyéndose el hialoplasma y los orgamelos de un modo equitativo.
Cuando no ocurre citocinesis luego de la caruccinesis, los dos núcleos quedan en el mismo citoplasma y resulta una
célula binucleada.
División en células vegetales:
No hay centríolos ni ásteres pero se organiza el huso acromático.
Citocinesis: el citoplasma se divide mediante un tabique, que se forma por la agrupación de microtóbulos y vescículas. Las vescículas crecen, se ordenan y se funden entre sí originando la placa celular. Finalmente se arman las paredes celulares a partir de celulosa, hemicelulosa y pectina.
Meiosis
Es un
proceso de reducción cromática por el que los cromosomas se reducen a la mitad. En la meiosis I (etapa reduccionaria) se reduce el número diploide de cromosomas a la mitad (haploide) pero aún los cromosomas son dobles. En la meiosis II (etapa ecuacional) se mantiene el número cromosómico haploide conseguido en la etapa anterior. Los cromosomas son simples.
Meiosis I: Está precedida por una interfase durante la cual se duplica el materialo genético.
PROFASE I: La envoltura nuclear y el nucleolo se desorganizan, los centríolos migran a polos oppuestos, duplicándose y se ordena el huso acromáticop. Se divide en 5 etapas: Leptonema, Cigonema, Paquinema, Diplonema y Diacinesis.
PROMETAFASE I: Los cromosomas migran al plano ecuatorial de
la célula.
METAFASE I: Los cromosomas se alinean en el plano ecuatorial. Los 2 cromosomas del bivalente se unen por medio del centrómero a la misma fibra del uso acromático.
ANAFASE I: Los 2 cromosomas homólogos unidos a la misma fibra dek huso se repelen y migran a polos opuestos. Cada cromosoma está formado por 2 cromatimas.
TELOFASE I: Cuando los cromosomas llegaron a los polos, se desorganizan el huso acromático y los ásteres, se reprganizan la envoltura nuclear y los nucleolos y se constituyen los núcleos hijos.
Citocinesis: Se produce simultáneamentye con la telofase, y da como resultado 2
célula hijas con un número haploide de cromosomas.
Intercinesis: Es un período que tiene lugar entre la
meiosis I y II y no se realiza duplicación del ADN.
Meiosis II: Los
procesos de esta división son semejantes a los de una mitosis en una célula haploide.
PROFASE II: Se condensan los cromosomas, se desintegran los nucleolos, los centríolos migran a los polos y se duplican, formación del huso acromático y se desorganiza la envoltura nuclear.
PROMETAFASE II: Los cromosomas condensados migran a la placa ecuatorial de
la célula.
METAFASE II: Los cromosomas se alinean en la placa ecuatorial, y cada cromosoma se une a una fibra del huso acromático.
ANAFASE II: Se fusiona el centrómero y se separan las 2 cromátidas de cada cromosoma. Cada una migra a un polo diferente.
TELOFASE II: Los
grupos cromosómicos llegan a los polos, el huso acromático se desorganiza, se reorganizan la envoltura nuclear y el nucleolo, se dispersan los cromosomas y se transforman en cromatina.
Citocinesis: Separación de los citoplasmas de las células hijas.
El
proceso melótico parte de una célula diploide que da como resultado 2 haploides, y a partir de éstas dos (melosis II) se obtienen 4 haploides.
Melosis, variabilidad
genética y evolución
La reproducción sexual introduce una importante proporción de variaciones genéticas. Cuanto mayor sea la diversidad de gametas formadas en cada progenitor, mayor será la
probabilidad de originar combinaciones diferentes por fecundación, y mayor será la diversidad de los descendientes. Una célula diploide, con 2 pares de cromosomas homólogos, originará por melosis 4 gametas haploides (uno de la madre y otro del padre). En la Metafase I se va a determinar en qué sentido migrarán en la Anafase I. Hay dos opciones:
Puede ocurrir que los 2 cromosomas paternos migren juntos a un polo y los dos maternos al opuesto.
Puede ocurrir que migren al mismo polo el cromosoma materno del par homólogo y el paterno del par homólogo. Los otros cromosomas, migran al polo opuesto.
Trabajo realizado por:
Facundo López.
MESOGRAFÍA
LÓPEZ, Facundo. (fecha desconocida). "Fisiología celular, Meiosis y Mitosis". Consultado el 11 de agosto del 2007 de http://www.monografias.com/trabajos/fisiocelular/fisiocelular.shtml

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